uid=HFR,o=lter,dc=ecoinformatics,dc=org all public read doi:10.6073/pasta/a76844a44291a12d0a188eea953d27d7 David Foster https://orcid.org/0000-0003-1171-3762 David Orwig https://orcid.org/0000-0001-7822-3560 Audrey Barker Plotkin https://orcid.org/0000-0002-4473-9785 Researcher 2023 English

Hemlock (Tsuga canadensis) forests in New England are changing rapidly with the invasion of the hemlock woolly adelgid (HWA, Adelges tsugae), a non-native insect pest that kills hemlock trees. As the adelgid invades hemlock stands at Harvard Forest, we are monitoring how forest structure and function changes. We surveyed community vegetation structure and composition prior to adelgid infestation, and are repeating this survey as infestation progresses. These measurements will allow us to link the long-term history of these hemlock stands to other current studies of hemlock response to adelgid infestation.

hemlock woolly adelgid hemlock understory vegetation vegetation dynamics LTER controlled vocabulary disturbance LTER core area Harvard Forest HFR LTER USA HFR default

This dataset is released to the public under Creative Commons CC0 1.0 (No Rights Reserved). Please keep the dataset creators informed of any plans to use the dataset. Consultation with the original investigators is strongly encouraged. Publications and data products that make use of the dataset should include proper acknowledgement.

Creative Commons Zero v1.0 Universal https://spdx.org/licenses/CC0-1.0.html CC0-1.0 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf048 Prospect Hill Tract (Harvard Forest), Slab City Tract (Harvard Forest). Coordinates based on WGS84 datum. -72.22 -72.18 +42.55 +42.45 250 350 meter 2002 2022 genus Tsuga species canadensis eastern hemlock ongoing Information Manager Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 hf-im@lists.fas.harvard.edu Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 (978) 724-3595 https://harvardforest.fas.harvard.edu

In 2002, we surveyed understory vegetation in existing hemlock-dominated plots at the Harvard Forest. Three plots were in or adjacent to the Hemlock Mapped Tree Plot (HF031), three were plots established in 1996 by Jennifer Jenkins (Jenkins, J. C., Aber, J. D., Canham, C. 1999. Hemlock woolly adelgid impacts on community structure and N cycling rates in eastern hemlock forest. Canadian Journal of Forest Research 29: 630-645.), and four were plots established in 1995 by Jason McLachlan (HF029 and HF053). In each plot, we established two transects running north-south. Five 1 m2 subplots were spaced evenly along each transect. We labeled the NW corner, unless otherwise noted. Transects are marked with string and yellow tent stakes (no string at Jenkins plots), and plots are marked with pink flags. Within each sub-plot, we estimated percent cover for herbs, shrubs, and seedlings to the nearest one percent. We counted the number of seedlings by species. A seedling is a tree less than 1.3m tall. If rocks or coarse woody debris covered at least 5% of the plot, their percent cover was recorded. In addition, the depth of the organic soil layer was measured to the nearest 0.5 cm at the NW corner of each subplot. We searched each plot and compiled a species list for the entire 20m x 20m or 30m x 30m area. Saplings were tallied by species across the plot. A sapling is a tree greater than 1.3m tall but less than 8cm in diameter. For the Hemlock History plots, that means that some stems were counted both as saplings and trees, since the Hemlock History Plots had trees tagged down to 2.5cm dbh. They were noted live or dead (only live saplings counted in 2022). In 2012, we resampled the understory vegetation (seedlings, saplings, herbs, shrubs, species list, substrates) only in the four plots established by Jason McLachlan. Organic layer depth was not measured in 2012. In 2016, we resampled understory vegetation and organic layer depth in the three plots associated with the Hemlock Mapped Tree Plot, and the four McLachlan plots. In 2022, we sampled understory vegetation in the four McLachlan plots. We discontinued measurements in the three Jenkins plots because these had less long-term ecological data, and are located nearby and in very similar forest to one of the McLachlan plots.

Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 (978) 724-3595 https://harvardforest.fas.harvard.edu https://ror.org/059cpzx98 pointOfContact The Harvard Forest Long-Term Ecological Research (LTER) program examines ecological dynamics in the New England region resulting from natural disturbances, environmental change, and human impacts. National Science Foundation LTER grants: DEB-8811764, DEB-9411975, DEB-0080592, DEB-0620443, DEB-1237491, DEB-1832210. hf048-01-o-layer.csv o-layer hf048-01-o-layer.csv 2541 67908f9ec6b8245ee4ed99a7eda73852 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-01-o-layer.csv plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value year year of measurement YYYY 1 year NA missing value o.layer depth of organic soil layer to the nearest 0.5 cm centimeter 0.1 real NA missing value 170 hf048-02-substrate.csv substrate hf048-02-substrate.csv 4665 7e36c8f5704201ac8ae374559b45f4b9 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-02-substrate.csv plot plot code HG1 : 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value year year of measurement YYYY 1 year NA missing value rock percent cover of rocks (except for 2002, estimated to the nearest 5%) dimensionless 1 whole NA missing value tree percent cover of tree stem (except for 2002, estimated to the nearest 5%) dimensionless 1 whole NA missing value down.wood percent cover of down wood (except for 2002, estimated to the nearest 5%) dimensionless 1 whole NA missing value 250 hf048-03-herb.csv herb hf048-03-herb.csv 13844 8b1611e59f79ff42ef19d49816034ab3 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-03-herb.csv year year of measurement YYYY 1 year NA missing value plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value aranud percent cover of Aralia nudicaulis. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value aritri percent cover of Arisaema triphyllum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value clibor percent cover of Clintonia borealis. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value coptri percent cover of Coptis trifolia. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value denpun percent cover of Dennstaedtia punctilobula. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value dryspi percent cover of Dryopteris spinulosa. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value epirep percent cover of Epigaea repens. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value galspe percent cover of Galium species. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value hupluc percent cover of Huperzia lucidula. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value maican percent cover of Maianthemum canadense. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value medvir percent cover of Medeola virginiana. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value monuni percent cover of Monotropa uniflora. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value osmcin percent cover of Osmunda cinnanoneum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value oxaspe percent cover of Oxalis species. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value lysbor percent cover of Lysimachia borealis. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value triund percent cover of Trillium undulatum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value uvuses percent cover of Uvularia sessilifolia. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value unk percent cover of unknown species. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value carspe percent cover of Carex (to genus only). Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value gootes percent cover of Goodyera tesselata. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value lycuni percent cover of Lycopus uniflorus. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value 250 hf048-04-shrub.csv shrub hf048-04-shrub.csv 7234 04dc5d7286716f2b9ffc53e19224cbae 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-04-shrub.csv year year of measurement YYYY 1 year NA missing value plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value gaupro percent cover of Gaultheria procumbens. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value mitrep percent cover of Mitchella repens. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value ilemuc percent cover of Ilex mucronata. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value rhopri percent cover of Rhododendron periclymenoides. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value vacang percent cover of Vaccinium angustifolium. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value viblan percent cover of Viburnum lantanoides. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value vaccor percent cover of Vaccinium corymbosum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value unk percent cover of an unknown shrub. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value 250 hf048-05-seedling-density.csv seedling density hf048-05-seedling-density.csv 9246 2de997c0a656d86706c87da35d9c0db9 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-05-seedling-density.csv year year of measurement YYYY 1 year NA missing value plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value acru number of Acer rubrum seedlings observed in the 1m2 plot number 1 whole NA missing value bele.beal number of Betula lenta or Betula alleghaniensis (difficult to distinguish as seedlings) seedlings observed in the 1m2 plot number 1 whole NA missing value bepa number of Betula papyrifera seedlings observed in the 1m2 plot number 1 whole NA missing value fagr number of Fagus grandifolis seedlings observed in the 1m2 plot number 1 whole NA missing value nysy number of Nyssa sylvatica seedlings observed in the 1m2 plot number 1 whole NA missing value piru number of Picea rubens seedlings observed in the 1m2 plot number 1 whole NA missing value pist number of Pinus strobus seedlings observed in the 1m2 plot number 1 whole NA missing value prse number of Prunus serotina seedlings observed in the 1m2 plot number 1 whole NA missing value quru number of Quercus rubra seedlings observed in the 1m2 plot number 1 whole NA missing value tsca number of Tsuga canadensis seedlings observed in the 1m2 plot number 1 whole NA missing value unk number of seedlings of unknown species observed in the 1m2 plot dimensionless 1 whole NA missing value acpe number of seedlings of Acer pensylvanicum observed in the 1m2 plot dimensionless 1 whole NA missing value 250 hf048-06-seedling-cover.csv seedling cover hf048-06-seedling-cover.csv 9826 66a3eb7f68fb365f7c356a324f5a1406 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-06-seedling-cover.csv year year of measurement YYYY 1 year NA missing value plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 NA missing value subplot number of 1m2 subplot number of 1m2 subplot NA missing value acru percent cover of Acer rubrum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value bele.beal percent cover of Betula lenta or Betula alleghaniensis (difficult to distinguish as seedlings). Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value bepa percent cover of Betula papyrifera. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value fagr percent cover of Fagus grandifolis. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value nysy percent cover of Nyssa sylvatica. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value piru percent cover of Picea rubens. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value pist percent cover of Pinus strobus. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value prse percent cover of Prunus serotina. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value quru percent cover of Quercus rubra. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value tsca percent cover of Tsuga canadensis. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value unk percent cover of unknown species. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value acpe percent cover of Acer pensylvanicum. Cover was recorded to the nearest 1%. A cover % of 0.5% was assigned to species present within a subplot but with less than 1% cover. dimensionless 0.1 real NA missing value 250 hf048-07-sapling.csv sapling hf048-07-sapling.csv 2287 c7f8e3cda737f98fa01f7ea623ad7483 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-07-sapling.csv year year of measurement YYYY 1 year NA missing value plot plot code HG1 20m x 20m plot within hemlock woodlot mapped plot on Prospect Hill Tract, Compartment 2 (the one with the huge O layer closest to BG swamp) HG2 20m x 20m plot within hemlock woodlot mapped plot (has lots of hemlock seedlings) HG3 20m x 20m plot just outside hemlock woodlot PH2 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 2 PH7 30m x 30m hemlock history plot located on Prospect Hill Tract, Compartment 7 SC1 20m x 20m plot hemlock plot on Slab City Tract, Compartment 9 SC10 30m x 30m hemlock history plot located on Slab City Tract, Compartment 10 SC2 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC3 20m x 20m plot Jenkins plot on Slab City Tract, Compartment 9 SC9 30m x 30m hemlock history plot located on Slab City Tract, Compartment 9 NA missing value species species ACPE Acer pensylvanicum ACRU Acer rubrum BELE Betula lenta CADE Castanea dentata FAGR Fagus grandifolia FRNI Fraxinus nigra TSCA Tsuga canadensis NA missing value status status L live D dead (dead saplings not counted in 2022) NA missing value hwa hemlock woolly adelgid (measured on TSCA only, based on visual inspection of branches within reach) 1 present 0 absent NA missing value number number of stems in the 20x20 or 30x30 m plot number 1 whole NA missing value plot.size size of plot hectare 0.01 real NA missing value num.ha number of stems on a per hectare basis number 0.1 real NA missing value note note note NA missing value 62 hf048-08-species.csv species distribution hf048-08-species.csv 7449 72e4fcc1ba25848bb4f5ac2ce377f547 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p04/hf048/hf048-08-species.csv type vegetation type F ferns / lycopods G graminoids H herbs S shrubs T trees NA missing value name.Haines.2011 species name (Latin) species name (Latin) NA missing value old.name older Latin name used in past surveys older Latin name used in past surveys NA missing value common.name common name of the plant common name of the plant NA missing value hg1.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value hg2.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value hg3.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value ph2.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value ph7.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value sc9.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value sc10.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value sc1.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value sc2.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value sc3.02 presence or absence of each species in the plot in 2002 0 absent 1 present NA missing value ph2.12 presence or absence of each species in the plot in 2012 0 absent 1 present NA missing value ph7.12 presence or absence of each species in the plot in 2012 0 absent 1 present NA missing value sc9.12 presence or absence of each species in the plot in 2012 0 absent 1 present NA missing value sc10.12 presence or absence of each species in the plot in 2012 0 absent 1 present NA missing value hg1.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value hg2.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value hg3.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value ph2.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value ph7.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value sc9.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value sc10.16 presence or absence of each species in the plot in 2016 0 absent 1 present NA missing value ph2.22 presence or absence of each species in the plot in 2022 0 absent 1 present NA missing value ph7.22 presence or absence of each species in the plot in 2022 0 absent 1 present NA missing value sc9.22 presence or absence of each species in the plot in 2022 0 absent 1 present NA missing value sc10.22 presence or absence of each species in the plot in 2022 0 absent 1 present NA missing value 77 plot community long-term measurement https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf083 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf053 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf031 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf106 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf029