uid=HFR,o=lter,dc=ecoinformatics,dc=org all public read doi:10.6073/pasta/ce368557a84cdfc718064c822b7033fe Aaron Ellison https://orcid.org/0000-0003-4151-6081 Nicholas Gotelli https://orcid.org/0000-0002-5409-7456 2023 English

Ecologists have long recognized that coexistence of interacting species depends on a diversity of separate, but linked habitats. Most models of species coexistence treat these habitat "patches" as unchanging elements of the landscape, but in reality, they are constantly changing in size and shape. The structure and dynamics of the food webs that inhabit them both control and depend on these changes. In this five-year (2006-2011) collaborative project, we are using a combination of extensive field surveys and intensive field experiments to understand how species interactions change the spatial distribution of dynamic habitat patches across the landscape, and to determine how food webs are structured within and among these patches. The experimental system for this five-year research project is the Sarracenia "microecosystem": the northern pitcher-plant Sarracenia purpurea; the unique food web of bacteria, protozoa, rotifers, mites, and fly larvae that live within its rainwater-filled leaves; three species of bog-inhabiting ants that are the primary prey for this carnivorous plant; and larvae of the pitcher-plant moth, Exyra fax, which cut and drain pitchers and remove food web habitat. Results from surveys of bogs in all six New England state and field experiments conducted at bogs at Harvard Forest and Belchertown, Massachusetts will be used to parameterize a simulation model to predict changes in spatial structure of ant, moth, and plant populations, and the structure and composition of the aquatic food web associated with the plant. This work is funded by the U.S. National Science Foundation, through collaborative awards 0541680 and 0541936.

biogeography carnivorous plants food webs insects LTER controlled vocabulary populations LTER core area Harvard Forest HFR LTER USA HFR default

This dataset is released to the public under Creative Commons CC0 1.0 (No Rights Reserved). Please keep the dataset creators informed of any plans to use the dataset. Consultation with the original investigators is strongly encouraged. Publications and data products that make use of the dataset should include proper acknowledgement.

Creative Commons Zero v1.0 Universal https://spdx.org/licenses/CC0-1.0.html CC0-1.0 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf159 Tom Swamp Tract (Harvard Forest), Swift River Bog (Belchertown MA). Coordinates based on WGS84 datum. -74.3 -66.9 +47.5 +40.5 0 1900 meter 2007 2013 genus Dolichoderus species pustulatus genus Exyra species fax genus Formica species subaenescens genus Lasius species minutus genus Myrmica species lobifrons genus Papaipema species appassionata genus Sarracenia species purpurea genus Tapinoma species sessile complete Information Manager Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 hf-im@lists.fas.harvard.edu Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 (978) 724-3595 https://harvardforest.fas.harvard.edu

Bogs for extensive field sampling are identified in collaboration with state officials, natural heritage programs, conservation agencies, etc. Necessary permits are obtained from appropriate permitting authorities. At each bog, we establish a 10m x 10m grid in the pitcher plant zone (plot dimensions are altered if the habitat zone for pitcher plants is too narrow. The plot is gridded at 1-m intervals. At each flagged point, we record relative moisture level of the bog mat as (high and dry, low and damp, or flooded). We also search for ant workers or ant nests within 20-cm of each flag. Representative samples of workers are hand-collected and fixed in ethanol for later identification. Voucher specimens are stored in Harvard’s Museum of Comparative Zoology. From each grid point, we locate the nearest pitcher plant and measure the distance to it with a tape measure and record the compass direction. We then count the number of old pitchers, measure the length of the 1st new pitcher, the rosette diameter, and the flowering status (flower, fruit, aborted bud) on each of these neighbor plants. We will also note any records of damage to pitcher plants from herbivorous moth larvae. We then collect five 50 ml water samples, one from each corner of the plot and one from the center of the grid. Water samples are collected by pressing a plastic centrifuge tube into the bog mat until it fills with pore water. Samples are pooled, filtered, and analyzed for anions and cations. Concurrent with water collections, we also collect 5 small handfuls of Sphagnum moss; identifications of Sphagnum species are done in the lab and confirmed by taxonomic specialists. Voucher specimens are stored in Harvard’s Farlow Herbarium.

(1) Tom Swamp Experiment: In April 2007, we established four 6 x 6 m plots on the bog mat. In these plots, we manipulated pitcher-plant density and pitcher-plant spatial arrangement by transplanting 120 greenhouse-grown pitcher plants into each plot. Transplants are in 4" plastic pots and were placed in regular arrays within the peat mat. At the end of the experiment, all transplants and pots will be removed (destructive harvest). If any transplants flower, flowers are removed so that no seeds from non-Tom Swamp lines can establish in the field. On each transplant we measure: presence/absence of pitcher plant moths (July), food web structure in pitcher plants (end of experiment), occurrence and distribution of other pitcher plants (seedlings, nonflowering adult plants, adult plants) within 5 replicate 6 x 6 m plots (May, September), and growth rates of pitcher plants (September). (2) Belchertown experiment: In this experiment, we simulate herbivory by Exyra fax larvae cutting a hole in a Sarracenia leaf that mimics the effects of larval feeding. To increase the realism of the experiment, we rub the inside of the wound with scrapings from real Exyra attacks, to facilitate the transfer of fungi and bacteria that are often associated with natural bore holes. Four experimental treatments were applied to 80 randomly selected adult plants (at least 10 cm longest leaf) at Swift River Bog in Belchertown, Massachusetts: 1) control (no plant manipulations), 2) single Exyra attack (only the first leaf of the season has a hole-punch), 3) multiple Exyra attacks (all leaves are perforated with a hole-punch), 4) single leaf removal (the first leaf of the season is entirely removed before it opens). Treatments were applied in 2007 and 2009. Plant growth, prey capture, and ant-nest distributions were measured in July (ants) and September (growth, prey capture) annually from 2007-2010.

In data file hf159-05-MAPP-exp1-growth.csv, an entry of -1 in any field indicates that the plant was dead at that census.

Harvard Forest

324 North Main Street Petersham MA 01366 USA

(978) 724-3302 (978) 724-3595 https://harvardforest.fas.harvard.edu https://ror.org/059cpzx98 pointOfContact The Harvard Forest Long-Term Ecological Research (LTER) program examines ecological dynamics in the New England region resulting from natural disturbances, environmental change, and human impacts. National Science Foundation LTER grants: DEB-8811764, DEB-9411975, DEB-0080592, DEB-0620443, DEB-1237491, DEB-1832210. hf159-01-bog-chars.csv bog characteristics hf159-01-bog-chars.csv 8928 2160bda4f921d6e838742c75c36b1c0f 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-01-bog-chars.csv state one of six New England states CT Connecticut MA Massachusetts NH New Hampshire RI Rhode Island VT Vermont ME Maine bog letter code uniquely identifying each sampled bog letter code uniquely identifying each sampled bog name local name of the sampled bog local name of the sampled bog date date bog was sampled YYYY-MM-DD plotortn orientation of the y-axis of the plot (from point 1000,1000 to point 1000,10000), in degrees east of magnetic north degree 1 whole NA missing value lat latitude of SW corner (point 1000, 1000) of sampled plot in decimal degrees, to nearest 100th degree 0.01 real NA missing value long longitude of SW corner (point 1000, 1000) of sampled plot in decimal degrees, to nearest 100th degree 0.01 real NA missing value elevation elevation in meters above sea level meter 1 whole NA missing value nplants number of Sarracenia purpurea plants measured in the plot (maximum = 100; NA = no plants measured, only at Bowdich Reservoir Bogs, RI) number 1 natural NA missing value papaipema number of plants attacked by Papaipema appassionata number 1 whole NA missing value exyra number of plants with leaves exhibiting damage by Exyra fax number 1 whole NA missing value endothenia number of plants attacked by Endothenia deakana number 1 whole NA missing value bud number of plants that attempted to flower number 1 whole NA missing value flower number of plants that successfully flowered. Less than or equal to bud number 1 whole NA missing value fruit number of plants that successfully fruited. Less than or equal to flower number 1 whole NA missing value na concentration of Sodium measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value nh4 concentration of Ammonium measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value k concentration of Potassium measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value mg concentration of Magnesium measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value ca concentration of Calcium measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value cl concentration of Chloride measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value no3 concentration of Nitrate measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value po4 concentration of Phosphate measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value so4 concentration of Sulfate measured in water samples taken from the plot. Values are from subsample of five pooled 50-ml samples taken at corners and center of plot. milligramsPerLiter 0.01 real NA missing value 78 hf159-02-bog-sarrs.csv bog sarracenia hf159-02-bog-sarrs.csv 623829 934b979c45c4a5bd58f5c1c9cb1c6b41 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-02-bog-sarrs.csv state one of the six New England states. See HF159-01-bog-chars.csv one of the six New England states bog code for sampled site. See HF159-01-bog-chars.csv for more detailed description of each bog code for sampled site grid.x location, in millimeters, of sample. Point (1000, 1000) is the SW corner of the plot, point (10000,10000) is the NE corner of the plot, and point (5000, 5000) is the center of the plot. At three bogs – Beeslick and Pauchaug in Connecticut, Bowdich Reservoir in RI – samples were collected haphazardly throughout the non-gridded sampling area. millimeter 1 natural NA missing value grid.y location, in millimeters, of sample. Point (1000, 1000) is the SW corner of the plot, point (10000,10000) is the NE corner of the plot, and point (5000, 5000) is the center of the plot. At three bogs – Beeslick and Pauchaug in Connecticut, Bowdich Reservoir in RI – samples were collected haphazardly throughout the non-gridded sampling area. millimeter 1 natural NA missing value moisture qualitative description of hummock/hollow topography at location (gridX, gridY) H high and dry, on a Sphagnum hummock L low and damp, in a Sphagnum hollow F flooded; standing water visible in a Sphagnum hollow plant.x actual location, in millimeters relative to SW corner of the plot of each Sarracenia purpurea closest to location (grid.x, grid.y) millimeter 1 natural NA missing value plant.y actual location, in millimeters relative to SW corner of the plot of each Sarracenia purpurea closest to location (grid.x, grid.y) millimeter 1 natural NA missing value d.to.xy distance, in millimeters from (plant.x, plant.y) to (grid.x, grid.y). This is measured if unique.plant = 1 and interpolated if unique.plant = 0. If plant.x = grid.x and plant.y = grid.y, then d.to.xy = 0 millimeter 1 real NA missing value unique.plant flag indicating whether the plant was actually measured (1), of if it was measured as the closest plant to another (gridX, gridY) (0). If unique.plant = 0, this means that the plant is “shared” with a point (gridX, gridY). For descriptions of individual populations, subset on unique.plant = 1. 1 plant was actually measured 0 plant was measured as the closest plant to another (gridX, gridY) npitchers number of pitchers produced in current year up to date of measurement number 0.1 real NA missing value nphyllodes number of phyllodia (flat pitchers) produced in current year up to date of measurement number 1 whole NA missing value longestleaf length, in millimeters, of the longest pitcher or phyllode millimeter 0.1 real NA missing value rosediam Largest diameter, in mm, of the rosette. If npitchers + nphyllodes = 1, then rosediam = 0. millimeter 0.1 real NA missing value bud attempted to flower or at least produced bud (1) or did not (0) 1 attempted to flower or at least produced bud 0 did not attempt to flower or produce bud flower plant at least flowered (1) or did not (e.g., bud aborted or eaten) (0). QA/QC check: if bud = 0, then flower must = 0. 1 plant at least flowered 0 plant did not flower fruit plant had fruited by sampling date (1) or did not (e.g., bud aborted, flower eaten) (0). QA/QC check: if flower = 0, then fruit must = 0. 1 plant had fruited by sampling date 0 plant had not fruited by sampling date papiapema rhizome bored by Papaipema appassionata and frass pyramid visible (1), or not (0) 1 rhizome bored by Papaipema appassionata and frass pyramid visible 0 rhizome not bored by Papaipema appassionata and frass pyramid not visible exyra at least one pitchers mined by Exyra fax (1), or not (0). QA/QC check: if npitchers = 0, then exyra must = 0. 1 at least one pitchers mined by Exyra fax 0 no pitchers mined by Exyra fax exdam number of pitchers mined by Exyra fax (range 0 to npitchers). QA/QC check: if exyra = 0, then exdam must = 0; if exyra = 1, then 1 ≤ exdam ≤ npitchers. number 1 whole NA missing value endothenia flower/fruit attacked by Endothenia deakana (1) or not (0). QA/QC check: If flower = 0, then endothenia must = 0. 1 flower/fruit attacked by Endothenia deakana 0 flower/fruit not attacked by Endothenia deakana antinplant an ant nest present in a drained, old pitcher (1) or not (0) 1 ant nest present in a drained, old pitcher 0 no ant nest present in a drained, old pitcher antatpoint an ant nest is present within a 10-cm circle around (gridX, gridY) 1 ant nest is present within a 10-cm circle around (gridX, gridY) 0 no ant nest is present within a 10-cm circle around (gridX, gridY) ant.substrate type of vegetation the the ant nest is in. Usually “sphagnum”, “sarracenia”, but occasionally others. QA/QC check: if antinplant AND antatpoint = 0, then antsubstrate = NA type of vegetation the the ant nest is in antsp1 species of ant collected (if applicable). Alphanumeric string length 6 or 7; see HF147 for translation of species code to full genus, species. species of ant collected antsp2 second species of ant collected (if applicable). Alphanumeric string length 6 or 7; see HF147 for translation of species code to full genus, species. second species of ant collected antsp3 species of ant collected (if applicable). Alphanumeric string length 6 or 7; see HF147 for translation of species code to full genus, species. species of ant collected (if applicable). Alphanumeric string length 6 or 7; see HF147 for translation of species code to full genus, species. sortcode record number. 1 – max. As of spring 2010, max(sortcode) = 5778. record number 7477 hf159-03-bog-sphag.csv bog sphagnum hf159-03-bog-sphag.csv 24693 4f9c2ae5fe13899200482949924976d6 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-03-bog-sphag.csv state one of six New England states. See HF159-01-bog-chars.csv one of six New England states bog code for sampled site. See HF159-01-bog-chars.csv for more detailed description of each bog code for sampled site grid.x location, in millimeters, of sample. Point (1000,1000) is the SW corner of the plot, point (10000,10000) is the NE corner of the plot, and point (5000, 5000) is the center of the plot. At three bogs – Beeslick and Pauchaug in Connecticut, Bowdich Reservoir in RI – samples were collected haphazardly throughout the non-gridded sampling area. millimeter 1 natural NA missing value grid.y location, in millimeters, of sample. Point (1000,1000) is the SW corner of the plot, point (10000,10000) is the NE corner of the plot, and point (5000, 5000) is the center of the plot. At three bogs – Beeslick and Pauchaug in Connecticut, Bowdich Reservoir in RI – samples were collected haphazardly throughout the non-gridded sampling area. millimeter 1 natural NA missing value affine species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot angustifolium species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot angermanicum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot bartlettianum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot capillifolium species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot centrale species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot compactum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot cuspidatum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot fallax species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot fimbriatum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot flavicomans species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot flexuosum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot fuscum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot magellanicum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot nemoreum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot pallustre species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot papillosum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot recurvum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot rubellum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot russowii species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot subtile species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot tenellum species presence in the particular sample at each site 1 present 0 absent NA no Sphagnum at that point in the plot 390 hf159-04-MAPPexp1-plotmap.csv MAPP experiment 1 plotmap hf159-04-MAPPexp1-plotmap.csv 9520 353887dbfa6ab49d71ad9736bc4f8ee0 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-04-MAPPexp1-plotmap.csv plot.num experimental plot number. Range 1 – 4. experimental plot number. Range 1 – 4. row row mid-point (y-coordinate), in meters. Range 0.5 – 5.5 meter 0.1 real NA missing value column column mid-point (x-coordinate), in meters. Range 0.5 – 5.5 meter 0.1 real NA missing value num.plants number of plants at location (column, row). Range 1 – 6 number 1 whole NA missing value treat.group size group of plants placed at location (column, row). See file HF159-05-MAPPexp1-growth.csv for initial sizes of each individual plant. A mean rosette diameter of plants is 27 mm B mean rosette diameter of plants is 55 mm C mean rosette diameter of plants is 77 mm D mean rosette diameter of plants is 99 mm E mean rosette diameter of plants is 121 mm F mean rosette diameter of plants is 139 mm plant.num identifier for each plant. Range 1 – 504. See HF159-05-MAPPexp1-growth.csv for growth and survivorship data of each individual plant. identifier for each plant 504 hf159-05-MAPPexp1-growth.csv MAPP experiment 1 growth hf159-05-MAPPexp1-growth.csv 88486 2cc0b371430164074f26861718e1afa0 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-05-MAPPexp1-growth.csv plot.num experimental plot number. Range 1 – 4. experimental plot number. Range 1 – 4. plant.num identifier for each plant. Range 1 – 504; corresponds with HF159-04-MAPPexp1-plotmap.csv identifier for each plant. Range 1 – 504 censored for survivorship analysis 0 plant has died at some point during the experiment 1 plant is still alive at the end of the experiment pitchers.init number of pitchers at the start of the experiment, April 2007. NOTE: an entry of -1 in any of the following fields indicates that the plant was dead at that census. number 1 whole NA missing value length.init length of longest pitcher at the start of the experiment, April 2007 millimeter 1 whole NA missing value rose.init rosette diameter at the start of the experiment, April 2007. If pitchers.init = 1 then rose.init = NA millimeter 1 whole NA missing value pitchers.2007 number of pitchers in September 2007; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2007 Number of phyllodia (flat pitchers) in September 2007; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value bud.2008 flower bud presence in May 2008 1 flower bud present 0 flower bud absent -1 plant was dead at that census flower.2008 flower presence in June 2008 1 flower present 0 flower absent -1 plant was dead at that census fruit.2008 fruit presence in September 2008 1 fruit present 0 fruit absent -1 plant was dead at that census exyra.2008 Exyra fax presence in July 2008 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2008 Papaipema appassionata presence in July 2008 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census ant.2008 if ant found nesting within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here. Otherwise NA. See HF147 for translation of species code to actual genus and species. An entry of -1 indicates that the plant was dead at that census if ant found nesting within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here pitchers.2008 number of pitchers in September 2008; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2008 number of phyllodes in September 2008; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value length.2008 length of longest pitcher in September 2008; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value rose.2008 rosette diameter in September 2008; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value bud.2009 flower bud presence in May 2009 1 flower bud present 0 flower bud absent -1 plant was dead at that census flower.2009 flower presence in June 2009 1 flower present 0 flower absent -1 plant was dead at that census fruit.2009 fruit presence in September 2009 1 fruit present 0 fruit absent -1 plant was dead at that census exyra.2009 Exyra fax presence in July 2009 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2009 Papaipema appassionata presence in July 2009 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census ant.2009 if ant found nesting within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here. See HF147 for translation of species code to actual genus and species. An entry of -1 indicates that the plant was dead at that census if ant found nesting within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here pitchers.2009 number of pitchers in September 2009; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2009 number of phyllodes in September 2009; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value length.2009 length of longest pitcher in September 2009; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value rose.2009 rosette diameter in September 2009; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value bud.2010 flower bud presence in May 2010 1 flower bud present 0 flower bud absent -1 plant was dead at that census flower.2010 flower presence in June 2010 1 flower present 0 flower absent -1 plant was dead at that census fruit.2010 fruit presence in September 2010 1 fruit present 0 fruit absent -1 plant was dead at that census exyra.2010 Exyra fax presence in July 2010 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2010 Papaipema appassionata presence in July 2010 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census ant.2010 if ant found nesting within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here. See HF147 for translation of species code to actual genus and species. An entry of -1 indicates that the plant was dead at that census six- or seven-character alphanumeric species code pitchers.2010 number of pitchers in September 2010; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2010 number of phyllodes in September 2010; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value length.2010 length of longest pitcher in September 2010; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value rose.2010 rosette diameter in September 2010; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value bud.2011 flower bud presence on 27 May 2011 1 flower bud present 0 flower bud absent -1 plant was dead at that census flower.2011 flower presence on 27 June 2011 1 flower present 0 flower absent -1 plant was dead at that census fruit.2011 fruit presence on 28 June 2011 1 fruit present 0 fruit absent -1 plant was dead at that census exyra.2011 Exyra fax presence on 28 June 2011 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2011 Papaipema appassionata presence on 28 June 2011 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census ant.2011 if ant found nesting on 28 June 2011 within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here. See HF147 for translation of species code to actual genus and species. An entry of -1 indicates that the plant was dead at that census six- or seven-character alphanumeric species code pitchers.2011 number of pitchers on 4 September 2011; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2011 number of phyllodes on 4 September 2011; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value length.2011 length of longest pitcher on 4 September 2011; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value rose.2011 rosette diameter on 4 September 2011; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value bud.2012 flower bud presence on 28 May 2012 1 flower bud present 0 flower bud absent -1 plant was dead at that census flower.2012 flower presence on 28 May 2012 1 flower present 0 flower absent -1 plant was dead at that census fruit.2012 fruit presence on 25 July 2012 1 fruit present 0 fruit absent -1 plant was dead at that census exyra.2012 Exyra fax presence on 25 July 2012 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2012 Papaipema appassionata presence on 25 July 2012 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census ant.2012 if ant found nesting on 25 July 2012 within 20-cm of plant, the six- or seven-character alphanumeric species code is entered here. See HF147 for translation of species code to actual genus and species. An entry of -1 indicates that the plant was dead at that census six- or seven-character alphanumeric species code pitchers.2012 number of pitchers on 4 September 2012; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2012 number of phyllodes on 4 September 2012; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value length.2012 length of longest pitcher on 4 September 2012; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value rose.2012 rosette diameter on 4 September 2012; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value pitchers.2013 number of pitchers on 28 August 2013; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value phyllodes.2013 number of phyllodes on August 28 2013; an entry of -1 indicates that the plant was dead at that census number 1 real NA missing value exyra.2013 Exyra fax presence on 28 August 2013 1 Exyra fax present 0 Exyra fax absent -1 plant was dead at that census papaipema.2013 Papaipema appassionata presence on 28 August 2013 1 Papaipema appassionata present 0 Papaipema appassionata absent -1 plant was dead at that census length.2013 length of longest pitcher on 28 August 2013; an entry of -1 indicates that the plant was dead at that census millimeter 1 real NA missing value shootmass.2013 oven-dried mass of aboveground tissue on 28 August 2013 gram 0.01 real NA missing value rootmass.2013 oven-dried mass of belowground tissue on 28 August 2013 gram 0.01 real NA missing value 504 hf159-06-MAPPexp3.csv MAPP experiment 3 hf159-06-MAPPexp3.csv 7553 da3c7948e853d6e8ae8c1e03b00c3dd7 1 \r\n column , https://harvardforest.fas.harvard.edu/data/p15/hf159/hf159-06-MAPPexp3.csv plant plant number. Range 501 – 580. Plants 570 and 575 do not exist. plant number treatment moth simulation treatment applied 1st CUT first leaf with hole punch in 2007 and 2009 to simulate Exyra damage ALL CUT all leaves with holes punched in 2007 and 2009 to simulate Exyra and Papaipema damage REMOVE first leaf removed in 2007 and 2009 control no treatment censored for survivorship analysis 0 plant has died at some point during the experiment 1 plant is still alive at the end of the experiment bud.2008 flower bud presence in May 2008. NOTE: an entry of -99999 in any of the following fields indicates that the plant was dead at that census. 1 flower bud present 0 flower bud absent flower.2008 flower presence in June 2008 1 flower present 0 flower absent fruit.2008 fruit presence in September 2008 1 fruit present 0 fruit absent ant.2008 if ant found nesting in July 2008 within 100-cm of plant, the six- or seven-character alphanumeric species code is entered here. Otherwise NA. See HF147 for translation of species code to actual genus and species. ant species code d.ant.2008 distance to ant nest from plant in July 2008 centimeter 1 whole NA missing value length.2008 length of longest pitcher in September 2008 millimeter 1 natural NA missing value rose.2008 rosette diameter in September 2008. If pitchers.2008 + phyllodes.2008 = 1, then rose.2008 = NA millimeter 1 natural NA missing value pitchers.2008 number of pitchers present in September 2008 number 1 whole NA missing value phyllodes.2008 number of phyllodes (flattened pitchers) present in September 2008 number 1 whole NA missing value bud.2009 flower bud presence in May 2009 1 flower bud present 0 flower bud absent flower.2009 flower presence in June 2009 1 flower present 0 flower absent fruit.2009 fruit presence in September 2009 1 fruit present 0 fruit absent ant.2009 if ant found nesting in July 2009 within 100-cm of plant, the six- or seven-character alphanumeric species code is entered here. Otherwise NA. See HF147 for translation of species code to actual genus and species. ant species code d.ant.2009 distance to ant nest from plant in July 2009 centimeter 1 whole NA missing value length.2009 length of longest pitcher in September 2009 millimeter 1 whole NA missing value rose.2009 rosette diameter in September 2009. If pitchers.2009 + phyllodes.2009 = 1, then rose.2009 = NA millimeter 1 whole NA missing value pitchers.2009 number of pitchers present in September 2009 number 1 whole NA missing value phyllodes.2009 number of phyllodes (flattened pitchers) present in September 2009 number 1 whole NA missing value bud.2010 flower bud presence in May 2010 1 flower bud present 0 flower bud absent flower.2010 flower presence in June 2010 1 flower present 0 flower absent fruit.2010 fruit presence in September 2010 1 fruit present 0 fruit absent ant.2010 if ant found nesting in July 2010 within 100-cm of plant, the six- or seven-character alphanumeric species code is entered here. Otherwise NA. See HF147 for translation of species code to actual genus and species. six- or seven-character alphanumeric species code d.ant.2010 distance to ant nest from plant in July 1010 centimeter 1 whole NA missing value length.2010 length of longest pitcher in September 2010 millimeter 0.1 real NA missing value rose.2010 rosette diameter in September 2010. If pitchers.2010 + phyllodes.2010 = 1, then rose.2010 = NA millimeter 1 whole NA missing value pitchers.2010 number of pitchers present in September 2010 number 1 whole NA missing value phyllodes.2010 number of phyllodes (flattened pitchers) present in September 2010 (number ). In 2010, always = 0 number 1 whole NA missing value pitchermass.2010 total dry mass of all pitchers harvested in September 2010 gram 0.01 real NA missing value 78 community short-term measurement https://harvardforest.fas.harvard.edu/ellison/current-research/spatial-dimension https://harvardforest.fas.harvard.edu/data/p15/hf159/HUH-10384.pdf https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf147 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf112 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf109 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf096 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf146 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf111 https://harvardforest.fas.harvard.edu/exist/apps/datasets/showData.html?id=hf114