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Frey, S

• Anthony, M. A., Frey, S. D., Stinson, K. A. 2017. Fungal community homogenization, shift in dominant trophic guild, and appearance of novel taxa with biotic invasion. Ecosphere 8: 17 pp.
• Anthony, M. A., Knorr, M., Moore, J. A. M., Simpson, M., Frey, S. D. 2021. Fungal community and functional responses to soil warming are greater than for soil nitrogen enrichment. Elementa: Science of the Anthropocene 9: article 000059, 15 pp.
• Anthony, M. A., Stinson, K. A., Moore, J. A. M., Frey, S. D. 2020. Plant invasion impacts on fungal community structure and function depend on soil warming and nitrogen enrichment. Oecologia 194: 659–672.
• Anthony, M. A., Stinson, K. A., Trautwig, A. N., Coates-Connor, E., Frey, S. D. 2019. Fungal communities do not recover after removing invasive Alliaria petiolata (garlic mustard). Biological Invasions 21: 3085-3099.
• Bradford, M. A., Davies, C. A., Frey, S. D., Maddox, T. R., Melillo, J. M., Mohan, J., Reynolds, J. F., Treseder, K. K., Wallenstein, M. D. 2008. Thermal adaptation of soil microbial respiration to elevated temperature. Ecology Letters 11: 1316-1327.
• Bradford, M. A., Wallenstein, M., Allison, S. D., Treseder, K. K., Frey, S., Watts, S. B., Davies, C. A., Maddox, T. R., Melillo, J. M., Mohan, J., Reynolds, J. F. 2009. Decreased mass specific respiration under experimental warming is robust to the microbial biomass method employed. Ecology Letters 12: E15-E18.
• Brangarí, A. C., Knorr, M. A., Frey, S. D., Rousk, J. 2025. Shifts in Microbial Thermal Traits Mitigate Heat-Induced Carbon Losses in Soils. Global Change Biology 31: e70579.
• Brzostek, E. R., Blair, J. M., Dukes, J. S., Frey, S. D., Hobbie, S. E., Melillo, J. M., Mitchell, R. J., Pendall, E., Reich, P. B., Shaver, G. R., Stefanski, A., Tjoelker, M. G., Finzi, A. C. 2012. The effect of experimental warming and precipitation change on proteolytic enzyme activity: positive feedbacks to nitrogen availability are not universal. Global Change Biology 18: 2617-2625.
• Burton, A. J., Melillo, J. M., Frey, S. D. 2008. Adjustment of Forest Ecosystem Root Respiration as Temperature Warms. Journal of Integrative Plant Biology 50: 1462-1483.
• Carey, J. C., Tang, J., Templer, P. H., Kroeger, K. D., Crowther, T. W., Burton, A. J., Dukes, J. S., Emmett, B., Frey, S. D., Heskel, M. A., Jiang, L., Machmuller, M. B., Mohan, J., Panetta, A., Reich, P. B., Reinsch, S., Wang, X., Allison, S. D. , Bamminger, C., Bridgham, S., Collins, S. L., de Dato, G., Eddy, W. C., Enquist, B. J., Estiarte, M., Harte, J., Henderson, A., Johnson, B. R., Steenberg Larsen, K., Luo, Y., Marhan, S., Melillo, J. M., Peñuelas, J., Pfeifer-Meister, L., Poll, C., Rastetter, E., Reinmann, A. B., Reynolds, L. L., Schmidt, I. K., Shaver, G. R., Strong, A. L., Suseela, V., Tietema, A. 2016. Temperature response of soil respiration largely unaltered with experimental warming. PNAS 113: 13797–13802.
• Chari, N. R., DeAngelis, K. M., Aguilar, A. A., Chan, A. L. H., Burgin, G. A., Frey, S. D., Taylor, B. N. 2026. Warming mitigates root exudate-induced priming effects via changes to microbial biomass, community structure, and gene abundance. The ISME Journal 20: wrag002.
• Chari, N. R., Muratore, T. J., Frey, S. D., Winters*, C. L., Martinez*, G., Taylor, B. N. 2024. Long-Term Soil Warming Drives Different Belowground Responses in Arbuscular Mycorrhizal and Ectomycorrhizal Trees. Global Change Biology 30: e17550.
• Conant, R. T., Ryan, M. G., Ågren, G. I., Birge, H. E., Bradford, M. A., Davidson, E. A., Eliasson, P. E., Evans, S. E., Frey, S. D., Giardina, C. P., Hopkins, F., Hyvönen, R., Kirschbaum, M. U. F., Lavallee, J. M., Leifeld, J., Parton, W. J., Steinweg, J. M., Wallenstein, M. D., Wetterstedt, J. A. M. 2011. Temperature and soil organic matter decomposition rates – synthesis of current knowledge and a way forward. Global Change Biology 17: 3392-3404.
• Contosta, A. R., Frey, S. D., Cooper, A. B. 2011. Seasonal dynamics of soil respiration and nitrogen mineralization in chronically warmed and fertilized soils. Ecosphere 2: 1-21.
• Contosta, A. R., Frey, S. D., Cooper, A. B. 2015. Soil microbial communities vary as much over time as with chronic warming and nitrogen additions. Soil Biology and Biochemistry 88: 19-24.
• Contosta, A. R., Frey, S. D., Ollinger, S. V., Cooper, A. 2013. Soil respiration does not acclimatize to warmer temperatures when modeled over seasonal timescales. Biogeochemistry 112: 555-570.
• Crowther, T. W., Thomas, S. M., Maynard, D. S., Baldrian, P., Covey, K., Frey, S. D., van Diepen, L. T. A., Bradford, M. A. 2015. Biotic interactions mediate soil microbial feedbacks to climate change. PNAS 112: 7033-7038.
• Crowther, T. W., Todd-Brown, K. E. O., Rowe, C. W., Wieder, W. R., Carey, J. C., Machmuller, M. B., Snoek, B. L., Fang, S., Zhou, G., Allison, S. D., Blair, J. M., Bridgham, S. D., Burton, A. J., Carrillo, Y., Reich, P. B., Clark, J. S., Classen, A. T., Dijkstra, F. A., Elberling, B., Emmett, B. A., Estiarte, M., Frey, S. D., Guo, J., Harte, J., Jiang, L., Johnson, B. R., Kröel-Dulay, G., Larsen, K. S., Laudon, H., Lavallee, J. M., Luo, Y., Lupascu, M., Ma, L. N., Marhan, S., Michelsen, A., Mohan, J., Niu, S., Pendall, E., Peñuelas, J., Pfeifer-Meister, L., Poll, C., Reinsch, S., Reynolds, L. L., Schmidt, I. K., Sistla, S., Sokol, N. W., Templer, P. H., Treseder, K. K., Welker, J. M., Bradford, M. A. 2016. Quantifying global soil carbon losses in response to warming. Nature 540: 104-108.
• DeAngelis, K. M., Pold, G., Topcuonglu, B. D., van Diepen, L. T. A., Varney, R., Blanchard, J., Melillo, J. M., Frey, S. D. 2015. Long-term forest soil warming alters microbial communities in temperate forest soils. Frontiers in Microbiology 6: 104.
• DeAngelis, K. M., Priyanka, R. C., Pold, G., Romero Olivares, A., Frey, S. D. 2019. Microbial responses to experimental soil warming: Five testable hypotheses. In: Mohan, J. E., Ecosystem Consequences of Soil Warming, Academic Press, Cambridge.
• Domeignoz-Horta, L. A., Pold, G., Erb, H., Sebag, D., Verrecchia, E., Northen, T., Louie, K., Eloe-Fadrosh, E., Pennacchio, C., Knorr, M. A., Frey, S. D., Melillo, J. M., DeAngelis, K. M. 2023. Substrate availability and not thermal acclimation controls microbial temperature sensitivity response to long-term warming. Global Change Biology 29: 1574 - 1590.
• Domeignoz-Horta, L. A., Pold, G., Liu, X. A., Frey, S. D., Melillo, J. M., DeAngelis, K. M. 2020. Microbial diversity drives carbon use efficiency in a model soil. Nature Communications 11: article 3684, 10 pp.
• Finzi, A. C., Ausin, A. T., Cleland, E. E., Frey, S. D., Houlton, B. Z., Wallenstein, M. D. 2011. Responses and feedbacks of coupled biogeochemical cycles to climate change: examples from terrestrial ecosystems. Frontiers in Ecology and the Environment 9: 61-67.
• Finzi, A. C., Giasson, M.-A., Barker Plotkin, A., Aber, J. D., Boose, E. R., Davidson, E. A., Dietze, M. C., Ellison, A. M., Frey, S. D., Goldman, E., Keenan, T. F., Melillo, J. M., Munger, J. W., Nadelhoffer, K. J., Ollinger, S. V., Orwig, D. A., Pederson, N., Richardson, A. D., Savage, K., Tang, J., Thompson, J. R., Williams, C. A., Wofsy, S. C., Zhou, Z., Foster, D. R. 2020. Carbon budget of the Harvard Forest Long-Term Ecological Research site: pattern, process, and response to global change. Ecological Monographs 90: e01423, 37 pp.
• Frey, S. D. 2006. Forest ecosystem responses to chronic nitrogen additions. Teaching Issues and Experiments in Ecology 4: 1-14.
• Frey, S. D. 2019. Mycorrhizal Fungi as Mediators of Soil Organic Matter Dynamics. Annual Review of Ecology, Evolution, and Systematics 50: 237-259.
• Frey, S. D., Drijber, R., Smith, H., Melillo, J. 2008. Microbial biomass, functional capacity, and community composition after twelve years of soil warming. Soil Biology and Biochemistry 40: 2904-2907.
• Frey, S. D., Knorr, M., Parrent, J. L., Simpson, R. T. 2004. Chronic nitrogen enrichment affects the structure and function of the soil microbial community in temperate hardwood and pine forests. Forest Ecology and Management 196: 159-171.
• Frey, S. D., Lee, J., Melillo, J. M., Six, J. W. 2013. The temperature response of soil microbial efficiency and its feedback to climate. Nature Climate Change 3: 395-398.
• Frey, S. D., Ollinger, S. V., Nadelhoffer, K. J., Bowden, R. D., Brzostek, E. R., Burton, A. J., Caldwell, B. A., Crow, S., Goodale, C. G., Grandy, A. S., Finzi, A. C., Kramer, M. G., Lajtha, K., LeMoine, J., Martin, M., McDowell, W. H., Minocha, R., Sadowsky, J. J., Templer, P. H., Wickings, K. 2014. Chronic nitrogen additions suppress decomposition and sequester soil carbon in temperate forests. Biogeochemistry 121: 305-316.
• Geyer, K. M., Schnecker, J., Grandy, A. S., Richter, A., Frey, S. D. 2020. Assessing microbial residues in soil as a potential carbon sink and moderator of carbon use efficiency. Biogeochemistry 151: 237–249.
• Giasson, M.-A., Ellison, A. M., Bowden, R. D., Crill, P. M., Davidson, E. A., Drake, J. E., Frey, S. D., Hadley, J. L., Lavine, M., Melillo, J. M., Munger, J. W., Nadelhoffer, K. J., Nicoll, L., Ollinger, S. V., Savage, K., Steudler, P. A., Tang, J., Varner, R. K., Wofsy, S. C., Foster, D. R., Finzi, A. C. 2013. Soil respiration in a northeastern US temperate forest: a 22-year synthesis. Ecosphere 4: article 140.
• Gilliam, F. S., Burns, D. A., Driscoll, C. T., Frey, S. D., Lovett, G. M., Watmough, S. A. 2019. Decreased atmospheric nitrogen deposition in eastern North America: Predicted responses of forest ecosystems. Environmental Pollution 244: 560-574.
• Haines, D. F., Aylward, J. A., Frey, S. D., Stinson, K. A. 2018. Regional Patterns of Floristic Diversity and Composition in Forests Invaded by Garlic Mustard (Alliaria petiolata). Northeastern Naturalist 25: 399-417.
• Harms, T. K., Groffman, P. M., Aluwihare, L., Craft, C., Wieder, W. R., Hobbie, S. E., Baer, S. G., Blair, J. M., Frey, S. D., Remucal, C. K., Rudgers, J. A., Collins, S. L. 2021. Patterns and trends of organic matter processing and transport: Insights from the US long-term ecological research network. Climate Change Ecology 2: article 100025, 12 pp.
• Heuck, C., Smolka, G., Whalen, E., Frey, S. D., Gundersen, P., Moldan, F., Fernandez, I. J., Spohn, M. 2018. Effects of long-term nitrogen addition on phosphorus cycling in organic soil horizons of temperate forests. Biogeochemistry 141: 167-181.
• Jones, J. A., Groffman, P. M., Blair, J., Davis, F. W., Dugan, H., Euskirchen, E. S., Frey, S. D., Harms, T. K., Hinckley, E., Kosmala, M., Loberg, S., Malone, S., Novick, K., Record, S., Rocha, A. V., Ruddell, B. L., Stanley, E. H., Sturtevant, C., Thorpe, A., White, T., Wieder, W. R., Zhai, L., Zhu, K. 2021. Synergies Among Environmental Science Research and Monitoring Networks: A Research Agenda. Earth's Future 9: e2020EF001631, 17 pp.
• Kamble, P. N., Rousk, J., Frey, S. D., Baath, E. 2013. Bacterial growth and growth limiting nutrients following chronic nitrogen additions to a hardwood forest soil. Soil Biology and Biochemistry 59: 32-37.
• Kenney, M. A., Kopp, R. E., Samaras, C., Alberti, M., Angell, S. Y., Burney, J., Cunliff, C., Duffy-Anderson, J. T., Fischer, Laura West, Frey, S. D., Hicke, J. A., Howard, P. H., Jerolleman, A., Kirchhoff, C. J., Koebele, E. A., Mahowald, N. M., Misarti, N., Pearson, D. L., Phillips, A. A., Siders, A. R., Singh, P. K., Smith, N. G., Teneva, L. T., Ulrich-Schad, J. D., Volenec, J. J., Akamani, K., Alexander, A. J., Anderson, C. E., Arbuckle, J. G., Auffhammer, M., Avery, C. W., Baker, S. H., Balk, D., Barbato, M., Barnard, P. L., Basaraba, A. B., Bednar, D. J., Bell, J. E., Bennett Gayle, D., Berry Daniels, L., Bertrand, D., Bistline, J. E. T., Brelsfoard, C. L., Briceno, T., Burkett, M., Burnham, M., Callahan, J. A., Cannon, C. E. B., Carothers, C., Carter, R. H., Cawley, M., Cha, J. M., Chang, H. H., Chang, M. H., Chegwidden, O. S., Chen, K., Cheng, B. S., Chi, G., Chu, E. K., Cleetus, R., Connor, A., Cook, A. A., Council, D., Cromar, K. R., Curtis, K. J., Daigle, J. J., DeLyser Roney, C., Domingue, S. J., Drake, J. E., Ebi, K. L., Elias, E. H., Fann, N. L., Fleishman, E., Gall, E. T., Ghitis, E., Gilmore, E. A., Gopalakrishnan, S., Grafton, D., Gray, S. A., Griffis, R. B., Grimm, N. B., Hadjimichael, A., Han, X., Haverkamp, J. A., Hess, J. J., Hessburg, P. F., Hinton, K. A., Hondula, D. M., Hubbell, B., Hultman, N., Jean, C., Jinnah, S., Johnson, J. X., Johnson, S. M., Johnson, V. A., Jones, K., Kaplan, P. O., Kay, S., Keenan, J. M., Keith, L., Kim, H. H., Kinney, P. L., Koller, S. F., Konfirst, M. A., Kosmal, A., Krosby, M. B., Lamp, W. O., Lanza, K., Lappen, J., Leach, J. E., Lee, C., Lehner, F., Li, Y., Luchinsky, L. A., Mach, K. J., Maldonado, J., Malone, S. L., Marchand, A. A. M., Marino, E., Markowitz, E. M., Masanet, E. R., Mauger, G. S., May, C. L., McConnell, K., McElwee, P., Mendez, M., Middel, A., Mladinich, S. K., Molina, R., Nassikas, N., Nateghi, R., Newman, R., O’Rourke, M. E., Obringer, R., Ortiz-Bobea, A., Osborne, E. B., Petersen, A., Preston, B. L., Puvvula, J., Raphael, M. N., Raymond, C. L., Reed, K. A., Ricke, K. L., Rising, J., Rocci, K. S., Rosenzweig, B. R., Ryder, S., Sanders, B. F., Sanderson, M. R., Schattman, R. E., Seeteram, N. A., Shi, L., Sidner, L., Siedlecki, S. A., Singh, D., Sonnerup, R. E., Steele, C., Tate, E., Thompson, J. R., Thompson, P. R., Turek-Hankins, L. L., Ulibarri, N., Ullrich, P. A., Wang, C., Ward, M., Ward Jones, M., Ward-Caviness, C., Whitehead, J. C., Wilson, M. T., Winkler, R. L., Wootten, A. M., Xian, G., Yorgey, G. G., York, A. M., Zuetell, E. J. 2026. Research Priorities for Robust Climate Assessments in the United States. Earth's Future 14: e2025EF007336.
• Knorr, M. A., Contosta, A. R., Morrison, E. W., Muratore, T. J., Anthony, M. A., Stoica, I., Geyer, K. M., Simpson, M. J., Frey, S. D. 2024. Unexpected sustained soil carbon flux in response to simultaneous warming and nitrogen enrichment compared with single factors alone. Nature Ecology & Evolution 10.1038/s41559-024-02546-x: 23 pp.
• Li, J., Wang, G., Mayes, M. A., Allison, S. D., Frey, S. D., Shi, Z., Hu, X., Luo, Y., Melillo, J. M. 2019. Reduced carbon use efficiency and increased microbial turnover with soil warming. Global Change Biology 25: 900-910.
• Liu, X. A., Frey, S. D., Melillo, J. M., DeAngelis, K. M. 2021. Physical protection regulates microbial thermal responses to chronic soil warming. Soil Biology and Biochemistry 159: article 108298, 5 pp.
• Liu, X. A., Pold, G., Domeignoz-Horta, L. A., Geyer, K. M., Caris, H., Nicholson, H., Kemner, K. M., Frey, S. D., Melillo, J. M., DeAngelis, K. M. 2021. Soil aggregate-mediated microbial responses to long-term warming. Soil Biology and Biochemistry 152: article 108055.
• Maynard, D. S., Covey, K. R., Crowther, T. W., Sokol, N. W., Morrison, E. W., Frey, S. D., Van Diepen, L. T. A., Bradford, M. A. 2018. Species associations overwhelm abiotic conditions to dictate the structure and function of wood-decay fungal communities. Ecology 99: 801-811.
• Melillo, J. M., Frey, S. D., DeAngelis, K. M., Werner, W. J., Bernard, M. J., Bowles, F. P., Pold, G., Knorr, M. A., Grandy, A. S. 2017. Long-term pattern and magnitude of soil carbon feedback to the climate system in a warming world. Science 358: 101-105.
• Moore, J. A. M., Anthony, M. A., Pec, G. J., Trocha, L. K., Trzebny, A., Geyer, K. M., van Diepen, L. T. A., Frey, S. D. 2021. Fungal community structure and function shifts with atmospheric nitrogen deposition. Global Change Biology 27: 1349-1364.
• Morrison, E. W., Frey, S. D., Sadowsky, J. J., van Diepen, L. T. A., Thomas, W. K., Pringle, A. 2016. Chronic nitrogen additions fundamentally restructure the soil fungal community in a temperate forest. Fungal Ecology 23: 48-57.
• Morrison, E. W., Pringle, A., Van Diepen, L. T. A., Frey, S. D. 2018. Simulated nitrogen deposition favors stress-tolerant fungi with low potential for decomposition. Soil Biology and Biochemistry 125: 75-85.
• Morrison, E. W., Pringle, A., Van Diepen, L. T. A., Grandy, A. S., Melillo, J. M., Frey, S. D. 2019. Warming alters fungal communities and litter chemistry with implications for soil carbon stocks. Soil Biology and Biochemistry 132: 120-130.
• Morrison, E. W., Whitney, S. A., Geyer, K. M., Sevigny, J. L., Grandy, A. S., Thomas, W. K., DeAngelis, K. M., Frey, S. D. 2022. Evidence for a genetic basis in functional trait tradeoffs with microbial growth rate but not growth yield. Soil Biology and Biochemistry 172: 108765.
• Muratore, T. J., Chari, N. R., Phillips, R. P., Taylor, B. N., Knorr, M. A., Frey, S. D. 2026. Increased root-derived carbon buffers soil carbon loss under simultaneous warming and nitrogen addition. Ecology 107: e70351.
• Muratore, T. J., Chari, N. R., Robinson, A.*, Gooijer, I.*, Welsh, S. G., Taylor, B. N., Frey, S. D., Tumber-Dávila, S. J. 2026. Coordination between root exudation rates and root conservation traits under simultaneous warming and nitrogen addition. New Phytologist 10.1111/nph.71398: .
• Muratore, T. J., Knorr, M. A., Simpson, M. J., Stephens, R. B., Phillips, R. P., Frey, S. D. 2024. Response of Root Respiration to Warming and Nitrogen Addition Depends on Tree Species. Global Change Biology 30: e17530.
• Pec, G. J., Van Diepen, L. T. A., Knorr, M., Grandy, A. S., Melillo, J. M., DeAngelis, K. M., Blanchard, J. L., Frey, S. D. 2021. Fungal community response to long-term soil warming with potential implications for soil carbon dynamics. Ecosphere 12: article e03460, 13 pp.
• Pisani, O., Frey, S. D., Simpson, A. J., Simpson, M. J. 2015. Soil warming and nitrogen deposition alter soil organic matter composition at the molecular-level. Biogeochemistry 123: 391-409.
• Pold, G., Billings, A. F., Blanchard, J. L., Burkhardt, D. B., Frey, S. D., Melillo, J. M., Schnabel, J., van Diepen, L. T. A., DeAngelis, K. M. 2016. Long-term warming alters carbohydrate degradation potential in temperate forest soils. Applied and Environmental Microbiology 82: 6518-6530.
• Pold, G., Domeignoz-Horta, L. A., Morrison, E. W., Frey, S. D., Sistla, S. A., DeAngelis, K. M. 2020. Carbon Use Efficiency and Its Temperature Sensitivity Covary in Soil Bacteria. mBio 11: e02293-19, 16 pp.
• Romero-Olivares, A. L., Davie-Martin, C. L., Kramshøj, M., Rinnan, R., Frey, S. D. 2022. Soil volatile organic compound emissions in response to soil warming and nitrogen deposition. Elementa: Science of the Anthropocene 10: 00065.
• Rousk, J., Frey, S. D. 2015. Revisiting the hypothesis that fungal-to-bacterial dominance characterises turnover of soil organic matter and nutrients. Ecological Monographs 85: 457-472.
• Rousk, J., Frey, S. D., Baath, E. 2012. Temperature adaptation of bacterial communities in experimentally warmed forest soils. Global Change Biology 18: 3252-3258.
• San Román, A. X., Frey, S. D., Knorr, M. A., Tong, H., Melillo, J. M., Simpson, M. J. 2026. Three decades of continuous warming in temperate forests destabilizes persistent forms of soil organic matter. Science of The Total Environment 1029: 181777.
• San Román, A. X., Srikanthan, N., Hamid, A. A., Muratore, T. J., Knorr, M. A., Frey, S. D., Simpson, M. J. 2024. Long-term warming in a temperate forest accelerates soil organic matter decomposition despite increased plant-derived inputs. Biogeochemistry 167: 1159-1174.
• Savage, K., Parton, W. J., Davidson, E. A., Trumbore, S. E., Frey, S. D. 2013. Long-term changes in forest carbon under temperature and nitrogen amendments in a temperate northern hardwood forest. Global Change Biology 19: 2389-2400.
• Shinfuku, M. S., Domeignoz-Horta, L. A., Choudoir, M. J., Frey, S.  D, Mitchell, M. F., Ranjan, R. 2024. Seasonal effects of long-term warming on ecosystem function and bacterial diversity. PLoS ONE 19: e0311364.
• Sierra, C. A., Trumbore, S. E., Davidson, E. A., Frey, S. D., Savage, K. E., Hopkins, F. M. 2012. Predicting decadal trends and transient responses of radiocarbon storage and fluxes in a temperate forest soil. Biogeosciences 9: 3013-3028.
• Sinsabaugh, R. L., Lauber, C. L., Weintraub, M. N., Ahmed, B., Allison, S. D., Crenshaw, C., Contosta, A. R., Cusack, D., Frey, S., Gallo, M. E., Gartner, T. B., Hobbie, S. E., Holland, K., Keeler, B. L., Powers, J. S., Stursova, C., Takacs-Vesbach, C., Waldrop, M., Wallenstein, M., Zak, D. R., Zeglin, L. H. 2008. Stoichiometry of soil enzyme activity at global scale. Ecology Letters 11: 1252-1264.
• Smith, M. D., La Pierre, K. L., Collins, S. L., Knapp, A. K., Gross, K. L., Barrett, J. E., Frey, S. D., Gough, L., Miller, R. B., Morris, J. T., Rustad, L. E., Yarie, J. 2015. Global environmental change and the nature of aboveground net primary productivity responses: Insights from long-term experiments. Oecologia 177: 935-947.
• Stinson, K. A., Frey, S. D., Jackson, M. R., Coates-Connor, E., Anthony, M., Martinez*, K. 2018. Responses of non-native earthworms to experimental eradication of garlic mustard and implications for native vegetation. Ecosphere 9, article e02353: 11 pp.
• Stoica, I., Tabatabaei Anaraki, M., Muratore, T., Knorr, M., Frey, S. D., Simpson, M. J. 2023. Chronic Warming and Nitrogen-Addition Alter Soil Organic Matter Molecular Composition Distinctly in Tandem Compared to Individual Stressors. ACS Earth and Space Chemistry 7: 609-622.
• Tonitto, C., Goodale, C. G., Weiss, M. S., Frey, S. D., Ollinger, S. V. 2014. The effect of nitrogen addition on soil organic matter dynamics: A model analysis of the Harvard Forest Chronic Nitrogen Amendment experiment and soil carbon response to anthropogenic N deposition. Biogeochemistry 117: 431-454.
• Van Diepen, L. T. A., Frey, S. D., Landis, E. A., Morrison, E. W., Pringle, A. 2017. Fungi exposed to chronic nitrogen enrichment are less able to decay leaf litter. Ecology 98: 5-11.
• Van Diepen, L. T. A., Frey, S. D., Sthultz, C. M., Morrison, E. W., Minocha, R., Pringle, A. 2015. Changes in litter quality caused by simulated nitrogen deposition reinforce the N-induced suppression of litter decay. Ecosphere 6: art205.
• Van de Broek, M., Riley, W. J., Tang, J., Frey, S. D., Schmidt, M. W. I. 2024. Thermal Adaptation of Enzyme-Mediated Processes Reduces Simulated Soil CO2 Fluxes Upon Soil Warming. Journal of Geophysical Research: Biogeosciences 129: e2024JG008619.
• VandenEnden, L., Anthony, M. A., Frey, S. D., Simpson, M. J. 2021. Biogeochemical evolution of soil organic matter composition after a decade of warming and nitrogen addition. Biogeochemistry 156: 161-175.
• Whalen, E. D., Smith, R. G., Grandy, A. S., Frey, S. D. 2018. Manganese limitation as a mechanism for reduced decomposition in soils under atmospheric nitrogen deposition. Soil Biology and Biochemistry 127: 252-263.
• Wheeler, J. A., Frey, S. D., Stinson, K. A. 2017. Tree seedling responses to multiple environmental stresses: Interactive effects of soil warming, nitrogen fertilization, and plant invasion. Forest Ecology and Management 403: 44-51.
• Wieder, W. R., Pierson, D., Earl, S., Lajtha, K., Baer, S. G., Ballantyne, S., Berhe, A. A., Billings, S. A., Brigham, L. M., Chacon, S. S., Fraterrigo, J., Frey, S. D., Georgiou, K, de Graaff, M., Grandy, A. S., Hartman, M. D., Hobbie, S. E., Johnson, C., Kaye, J., Kyker-Snowman, E., Litvak, M. E., Mack, M. C., Malhotra, A., Moore, J. A. M., Nadelhoffer, K., Rasmussen, C., Silver, W. L., Sulman. B. N., Walker. X., Weintraub, S. 2021. SoDaH: the SOils DAta Harmonization database, an open-source synthesis of soil data from research networks, version 1.0. Earth System Science Data 13: 1843–1854.
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